鳄鱼皮肤图案源自精确机械折叠
瑞士科学家研究显示,鳄鱼面部与下颌独特的鳞片图案是由一种精确的皮肤机械折叠过程形成的。研究结果验证了之前的观点,即这些鳞片来自某种机械过程而非遗传机制,同时为了解这一过程背后的细节提供了新的见解。相关研究12月12日发表于《自然》。
动物的附属器官——如羽毛、毛发和鳞片——通常作为受遗传调控的单元在胚胎发育期间发育。但这一规则也有例外,如鳄鱼头部的鳞片被发现完全由机械过程产生。不过,形成头部鳞片图案的精确机制一直很难确定,一定程度上是因为用鳄鱼胚胎做实验存在技术难度。
日内瓦大学的Michel Milinkovitch和同事通过将尼罗鳄胚胎实验与计算机模拟相结合,生成了一个三维机械发育模型,重现了鳄鱼头部鳞片的图案结构。他们发现,这些鳞片会通过简单的机械过程自我组织,比如因皮肤比皮下骨骼发育更快以及这些不同组织的硬度差异而导致的挤压褶皱。研究者总结道,这个过程会使鳄鱼头部的鳞片形成不规则的几何图案,这与鳄鱼身体鳞片作为受遗传调控单元的发育方式截然不同。(来源:中国科学报 冯维维)
幼年(2 岁)的尼罗鳄。图片来自:瑞士日内瓦大学A. Debry & M. C. Milinkovitch
相关论文信息:https://doi.org/10.1038/s41586-024-08268-1
版权声明:本文转载仅仅是出于传播信息的需要,并不意味着代表本网站观点或证实其内容的真实性;如其他媒体、网站或个人从本网站转载使用,须保留本网站注明的“来源”,并自负版权等法律责任;作者如果不希望被转载或者联系转载稿费等事宜,请与我们接洽。
已有 0条评论,共0赞
验证码: |
|
最新评论 |
|
element block and removes
// previous instances of the identical script.
var found = false;
for (var child = 0; child < head.childNodes.length; child++) {
var control = head.childNodes[child];
if (typeof(control.tagName) == "string") {
if (control.tagName.toUpperCase() == "SCRIPT") {
if (script.src.length > 0) {
if (script.src == control.src) {
found = true;
break;
}
} else if (script.innerHTML.length > 0) {
if (script.innerHTML == control.innerHTML) {
found = true;
break;
}
}
}
}
}
if (found) {
head.removeChild(control);
}
var scriptAddedToHead = false;
if (typeof script.readyState != "undefined" && !window.opera) {
script.onreadystatechange = function() {
if (script.readyState != "complete" && script.readyState != "loaded") {
return;
} else {
Anthem_LoadPageScript(result, index + 1);
}
}
} else {
if (isExternalScript) // if it's an external script, only execute the next script when the previous one is loaded.
{
script.onload = function() {
Anthem_LoadPageScript(result, index + 1);
}
}
else // I didn't find a way for script blocks to fire some onload event. So in this case directly call the Anthem_LoadPageScript for the next script.
{
document.getElementsByTagName('head')[0].appendChild(script);
scriptAddedToHead = true;
Anthem_LoadPageScript(result, index + 1);
}
}
// Now we append the new script and move on to the next script.
// Note that this is a recursive function. It stops when the
// index grows larger than the number of scripts.
if (!scriptAddedToHead)
document.getElementsByTagName('head')[0].appendChild(script);
}
} catch (e) {
Anthem_DebugError("Error adding page script to head. " + e.name + ": " + e.message);
}
}
}
function Anthem_EvalClientSideScript(result) {
if (result.script) {
for (var i = 0; i < result.script.length; ++i) {
try {
eval(result.script[i]);
} catch (e) {
alert("Error evaluating client-side script!\n\nScript: " + result.script[i] + "\n\nException: " + e);
}
}
}
}
//Fix for bug #1429412, "Reponse callback returns previous response after file push".
//see http://sourceforge.net/tracker/index.php?func=detail&aid=1429412&group_id=151897&atid=782464
function Anthem_Clear__EVENTTARGET() {
var form = Anthem_GetForm();
Anthem_SetHiddenInputValue(form, "__EVENTTARGET", "");
}
function Anthem_InvokePageMethod(methodName, args, clientCallBack, clientCallBackArg) {
Anthem_Clear__EVENTTARGET(); // fix for bug #1429412
return Anthem_CallBack(null, "Page", null, methodName, args, clientCallBack, clientCallBackArg, true, true);
}
function Anthem_InvokeMasterPageMethod(methodName, args, clientCallBack, clientCallBackArg) {
Anthem_Clear__EVENTTARGET(); // fix for bug #1429412
return Anthem_CallBack(null, "MasterPage", null, methodName, args, clientCallBack, clientCallBackArg, true, true);
}
function Anthem_InvokeControlMethod(id, methodName, args, clientCallBack, clientCallBackArg) {
Anthem_Clear__EVENTTARGET(); // fix for bug #1429412
return Anthem_CallBack(null, "Control", id, methodName, args, clientCallBack, clientCallBackArg, true, true);
}
function Anthem_PreProcessCallBack(
control,
e,
eventTarget,
causesValidation,
validationGroup,
imageUrlDuringCallBack,
textDuringCallBack,
enabledDuringCallBack,
preCallBackFunction,
callBackCancelledFunction,
preProcessOut
) {
var valid = true;
if (causesValidation && typeof(Page_ClientValidate) == "function") {
valid = Page_ClientValidate(validationGroup);
}
if (typeof(WebForm_OnSubmit) == "function") {
valid = WebForm_OnSubmit();
}
if (valid) {
var preCallBackResult = true;
if (typeof(preCallBackFunction) == "function") {
preCallBackResult = preCallBackFunction(control, e);
}
if (typeof(preCallBackResult) == "undefined" || preCallBackResult) {
var inputType = control.getAttribute("type");
inputType = (inputType == null) ? '' : inputType.toUpperCase();
if (inputType == "IMAGE" && e != null) {
var form = Anthem_GetForm();
if (e.offsetX) { // IE
Anthem_SetHiddenInputValue(form, eventTarget + ".x", e.offsetX);
Anthem_SetHiddenInputValue(form, eventTarget + ".y", e.offsetY);
} else { // FireFox + ???
var offset = GetControlLocation(control);
Anthem_SetHiddenInputValue(form, eventTarget + ".x", e.clientX - offset.x + 1 + window.pageXOffset);
Anthem_SetHiddenInputValue(form, eventTarget + ".y", e.clientY - offset.y + 1 + window.pageYOffset);
}
}
if (imageUrlDuringCallBack || textDuringCallBack) {
var nodeName = control.nodeName.toUpperCase();
if (nodeName == "INPUT") {
if (inputType == "CHECKBOX" || inputType == "RADIO" || inputType == "TEXT") {
preProcessOut.OriginalText = GetLabelText(control.id);
SetLabelText(control.id, textDuringCallBack);
} else if (inputType == "IMAGE") {
if (imageUrlDuringCallBack) {
preProcessOut.OriginalText = control.src;
control.src = imageUrlDuringCallBack;
} else {
preProcessOut.ParentElement = control.parentElement ? control.parentElement : control.parentNode;
if (preProcessOut.ParentElement) {
preProcessOut.OriginalText = preProcessOut.ParentElement.innerHTML;
preProcessOut.ParentElement.innerHTML = textDuringCallBack;
}
}
} else if (inputType == "SUBMIT" || inputType == "BUTTON") {
preProcessOut.OriginalText = control.value;
control.value = textDuringCallBack;
}
} else if (nodeName == "SELECT" || nodeName == "SPAN") {
preProcessOut.OriginalText = GetLabelText(control.id);
SetLabelText(control.id, textDuringCallBack);
} else {
preProcessOut.OriginalText = control.innerHTML;
control.innerHTML = textDuringCallBack;
}
}
// Disable the control during callback if required
control.disabled = (typeof(enabledDuringCallBack) == "undefined") ? false : !enabledDuringCallBack;
return true;
} else {
// Callback cancelled
if (typeof(callBackCancelledFunction) == "function") {
callBackCancelledFunction(control, e);
}
return false;
}
} else {
// Validation failed
return false;
}
}
function Anthem_PreProcessCallBackOut() {
// Fields
this.ParentElement = null;
this.OriginalText = '';
}
function Anthem_PostProcessCallBack(
result,
control,
e,
eventTarget,
clientCallBack,
clientCallBackArg,
imageUrlDuringCallBack,
textDuringCallBack,
postCallBackFunction,
preProcessOut
) {
if (typeof(postCallBackFunction) == "function") {
postCallBackFunction(control, e);
}
// Re-enable the control if it was disabled during callback
control.disabled = false;
var inputType = control.getAttribute("type");
inputType = (inputType == null) ? '' : inputType.toUpperCase();
if (inputType == "IMAGE") {
var form = Anthem_GetForm();
Anthem_RemoveHiddenInput(form, eventTarget + ".x");
Anthem_RemoveHiddenInput(form, eventTarget + ".y");
}
if (imageUrlDuringCallBack || textDuringCallBack) {
var nodeName = control.nodeName.toUpperCase();
if (nodeName == "INPUT") {
if (inputType == "CHECKBOX" || inputType == "RADIO" || inputType == "TEXT") {
SetLabelText(control.id, preProcessOut.OriginalText);
} else if (inputType == "IMAGE") {
if (imageUrlDuringCallBack) {
control.src = preProcessOut.OriginalText;
} else {
preProcessOut.ParentElement.innerHTML = preProcessOut.OriginalText;
}
} else if (inputType == "SUBMIT" || inputType == "BUTTON") {
control.value = preProcessOut.OriginalText;
}
} else if (nodeName == "SELECT" || nodeName == "SPAN") {
SetLabelText(control.id, preProcessOut.OriginalText);
} else {
control.innerHTML = preProcessOut.OriginalText;
}
}
if (typeof(clientCallBack) == "function") {
clientCallBack(result, clientCallBackArg);
}
}
function Anthem_FireCallBackEvent(
control,
e,
eventTarget,
eventArgument,
causesValidation,
validationGroup,
imageUrlDuringCallBack,
textDuringCallBack,
enabledDuringCallBack,
preCallBackFunction,
postCallBackFunction,
callBackCancelledFunction,
includeControlValuesWithCallBack,
updatePageAfterCallBack
) {
// Cancel the callback if the control is disabled. Although most controls will
// not raise their callback event if they are disabled, the LinkButton will.
// This check is for the LinkButton. See SourceForge Patch 1639700.
if (control.disabled) return;
var preProcessOut = new Anthem_PreProcessCallBackOut();
var preProcessResult = Anthem_PreProcessCallBack(
control,
e,
eventTarget,
causesValidation,
validationGroup,
imageUrlDuringCallBack,
textDuringCallBack,
enabledDuringCallBack,
preCallBackFunction,
callBackCancelledFunction,
preProcessOut
);
if (preProcessResult) {
var eventType = e.type;
Anthem_FireEvent(
eventTarget,
eventArgument,
function(result) {
Anthem_PostProcessCallBack(
result,
control,
eventType,
eventTarget,
null,
null,
imageUrlDuringCallBack,
textDuringCallBack,
postCallBackFunction,
preProcessOut
);
},
null,
includeControlValuesWithCallBack,
updatePageAfterCallBack
);
}
}
function AnthemListControl_OnClick(
e,
causesValidation,
validationGroup,
textDuringCallBack,
enabledDuringCallBack,
preCallBackFunction,
postCallBackFunction,
callBackCancelledFunction,
includeControlValuesWithCallBack,
updatePageAfterCallBack
) {
var target = e.target || e.srcElement;
if (target.nodeName.toUpperCase() == "LABEL" && target.htmlFor != '')
return;
var eventTarget = target.id.split("_").join("$");
Anthem_FireCallBackEvent(
target,
e,
eventTarget,
'',
causesValidation,
validationGroup,
'',
textDuringCallBack,
enabledDuringCallBack,
preCallBackFunction,
postCallBackFunction,
callBackCancelledFunction,
true,
true
);
}
// Returns the top, left control location in FireFox
function GetControlLocation(control) {
var offsetX = 0;
var offsetY = 0;
var parent;
for (parent = control; parent; parent = parent.offsetParent) {
if (parent.offsetLeft) {
offsetX += parent.offsetLeft;
}
if (parent.offsetTop) {
offsetY += parent.offsetTop;
}
}
return { x: offsetX, y: offsetY };
}
function GetLabelText(id) {
var labels = document.getElementsByTagName('label');
for (var i = 0; i < labels.length; i++) {
if (labels[i].htmlFor == id) {
return labels[i].innerHTML;
}
}
return null;
}
functio